pe
pep-10966 v1 CC-BY-SA-4.0

Dermaseptin S5: frog-skin natural germ-killer (DS5)

A natural peptide from South American leaf frog skin that kills bacteria, fungi, and parasites (including the one that causes Leishmania disease) by punching holes in their outer membranes; used only as a lab research tool.

statusdesigned target? length27 aa refs12
status 1 / 5
sequence27 aa
151015202527
ALWKTLLKK VGTHLAGKA ALGAAADKX
overview readme

What this is

Dermaseptin S5 (DS5, also Dermaseptin-S5 or DRS-S5) is a natural antimicrobial peptide found in the skin secretions of South American leaf frogs of the genus Phyllomedusa — the same family whose skin glands also produce the opioid peptide dermorphin (/card/pep-10694). It belongs to the dermaseptin family of cationic, lysine-rich peptides, roughly 27–34 residues long, that fold into an amphipathic α-helix on contact with microbial membranes and kill bacteria, fungi, and protozoan parasites by physically disrupting those membranes (Mor and colleagues, Biochemistry, 1994; Leite and colleagues, Comparative Biochemistry and Physiology Part A, 2008). DS5 is a preclinical research peptide: it has never been tested in humans, has no approved indication, and is studied primarily as a starting point for designing membrane-active anti-infectives. The stored sequence ALWKTLLKKVGTHLAGKAALGAAADKX (27 letters with a trailing X placeholder) represents the peptide backbone; the native peptide carries a C-terminal amide (-NH₂) cap that is not visible in the raw single-letter string and that protects the C-terminus from carboxypeptidases.

History

The dermaseptin family was first described in the early 1990s by the laboratories of Amram Mor and Pierre Nicolas, working on the skin secretions of Phyllomedusa tree frogs. The 1994 paper by Mor and colleagues in Biochemistry — "Structure, synthesis, and activity of Dermaseptin b, a novel vertebrate defensive peptide from frog skin: relationship with adenoregulin" — anchored the structural model that still defines the group: cationic, ~27–34 residue peptides that adopt an amphipathic α-helix on membranes. Two parallel research threads followed. Pierre and colleagues (European Journal of Biochemistry, 2000) and Chen and colleagues (Regulatory Peptides, 2003; Peptides, 2006) mapped out the precursor architecture, showing that the dermaseptin precursors share a conserved N-terminal preprosequence with the precursors of the opioid peptides dermorphin and the deltorphins — the same skin gland produces both families from related cDNAs. Vanhoye and colleagues (European Journal of Biochemistry, 2003) traced this precursor architecture back to a roughly 150-million-year-old ancestral gene with a conserved signal peptide but a hypermutable antimicrobial domain, which is why each Phyllomedusa species produces its own distinct dermaseptin cocktail.

DS5 itself sits within the S-series (S1–S5) from Phyllomedusa sauvagii. It has remained a reference compound for membrane-disruption biophysics and a comparator in structure–activity work on better-studied siblings such as dermaseptin S4 (Krugliak and colleagues, Antimicrobial Agents and Chemotherapy, 2000), rather than the lead of an independent drug-development program.

What it does

DS5 kills a broad range of microbes — Gram-positive and Gram-negative bacteria, yeasts and filamentous fungi, and protozoan parasites including Leishmania and Plasmodium — by tearing open their cell membranes (Leite and colleagues, 2008; Krugliak and colleagues, 2000; Ghosh and colleagues, Journal of Biological Chemistry, 1997). The peptide is largely disordered in water; on contact with the negatively charged outer surface of a microbial membrane it folds into an amphipathic α-helix, accumulates on the membrane, and either disorganizes the bilayer in a detergent-like "carpet" fashion or assembles into transient toroidal pores. Either way the membrane's electrochemical gradient collapses and the cell leaks ions and cytoplasmic contents.

Selectivity for microbes over host cells comes from chemistry, not specificity: bacterial and parasite membranes are richer in anionic lipids (phosphatidylglycerol, cardiolipin, lipopolysaccharide) and lack cholesterol, so the cationic peptide is drawn to them preferentially. That selectivity is relative, not absolute — at high enough concentrations dermaseptins damage mammalian cells too — and the therapeutic window is the central drug-development question for the whole family (Leite and colleagues, 2008). Because the mechanism is physical disruption of the membrane rather than inhibition of a specific enzyme, microbes have a hard time evading dermaseptins by point mutation; this is part of the long-running interest in the class as a response to antibiotic resistance.

Evidence

  • Human: No published human trials. There are no human pharmacokinetic, safety, or efficacy data for DS5 in any indication.
  • Animal: Limited. The native DS5 literature is dominated by in vitro assays; most in vivo work in the dermaseptin family has gone to engineered analogs of dermaseptin S4 (Krugliak and colleagues, 2000) rather than to DS5 itself.
  • In vitro: Broad antimicrobial activity at low-micromolar concentrations against Gram-positive and Gram-negative bacteria, Candida and filamentous fungi, Leishmania promastigotes, and intraerythrocytic Plasmodium falciparum (Ghosh and colleagues, 1997; Krugliak and colleagues, 2000; Leite and colleagues, 2008). DS5 is reported to have low hemolytic activity against human red blood cells at antimicrobial concentrations relative to several of its siblings (Leite and colleagues, 2008), which is part of why it has been used as a scaffold for analog design.

Known effects

  • Broad-spectrum antibacterial (Gram-positive and Gram-negative) — in vitro only
  • Antifungal activity against Candida — in vitro only
  • Anti-Leishmania activity (promastigotes) — in vitro; mechanism is direct parasite-membrane disruption
  • Anti-Plasmodium activity (intraerythrocytic stages) — in vitro; demonstrated more extensively for the S-series sibling dermaseptin S3 (Ghosh and colleagues, 1997) and for dermaseptin S4 derivatives (Krugliak and colleagues, 2000)
  • Lower hemolytic activity than several other dermaseptins at antimicrobial concentrations, in vitro (Leite and colleagues, 2008)

Safety signals

There is no human safety profile for DS5 — it has never been administered to humans in any published study. In vitro, DS5 is reported as having low hemolytic activity against human red blood cells at antimicrobial concentrations relative to several family members (Leite and colleagues, 2008), but membrane-disrupting peptides are cytotoxic to mammalian cells at high enough concentrations, and that ceiling is sequence- and context-dependent. The relative selectivity for microbial over mammalian membranes that DS5 shows in vitro is not the same thing as a characterized in vivo safety window.

Regulatory status

  • US: Not approved. No Investigational New Drug (IND) application on record. Research compound only.
  • EU: Not approved. No EMA marketing authorization or clinical trial authorization for any dermaseptin.
  • WADA: Not listed; not relevant as a research peptide with no human use pathway.

DS5 is a laboratory peptide; preparations from research-chemical channels are not authorized for human use and carry no identity, purity, sterility, or endotoxin guarantees.

Related peptides

  • Dermorphin (/card/pep-10694) — also from Phyllomedusa skin, but a μ-opioid receptor agonist rather than an antimicrobial peptide; encoded by precursors that share an N-terminal preprosequence with the dermaseptins (Pierre and colleagues, 2000; Vanhoye and colleagues, 2003).
  • LL-37 (/card/pep-00002) — the human cathelicidin antimicrobial peptide; a cationic amphipathic α-helix with the same membrane-disruption logic as the dermaseptins, but evolved independently in mammalian innate immunity.
  • Magainin — antimicrobial α-helix from the skin of the African clawed frog Xenopus laevis; the canonical example of an amphibian AMP outside the Phyllomedusa lineage.

Mechanism

DS5 is a cationic, lysine-rich peptide of approximately 28 residues that is largely unstructured in aqueous solution and adopts an amphipathic α-helical conformation upon contact with lipid bilayers. Electrostatic attraction to anionic phospholipid headgroups (phosphatidylglycerol, cardiolipin) and to lipopolysaccharide recruits the peptide to the outer leaflet of a microbial membrane, where the hydrophobic and hydrophilic faces of the helix align with the membrane environment. Peptide accumulates on the outer leaflet until a critical local density is reached, at which point it either disorganizes the bilayer in a "carpet"-like detergent fashion or assembles into transient toroidal pores — in either case the transmembrane gradient collapses, ions and small metabolites leak across, and the cell dies (Leite and colleagues, 2008). The same mechanism accounts for activity against bacteria, yeasts, Leishmania promastigotes, and intraerythrocytic Plasmodium stages, since all share negatively charged surface membranes that mammalian cell membranes do not.

Two further mechanism notes are worth flagging. First, dermaseptins in the native frog secretion appear to act as a cocktail rather than as individual agents — published work in the family has reported synergy between dermaseptin variants in combined assays, consistent with the hypothesis that the cocktail composition matters (Mor and colleagues, 1994). Second, the dermaseptin precursor architecture is the same one that generates the opioid peptides dermorphin and the deltorphins in the same skin glands (Pierre and colleagues, 2000; Vanhoye and colleagues, 2003), so the same animal lineage produces two pharmacologically unrelated peptide families from related cDNAs — a curiosity of amphibian skin biology that recurs throughout this literature.

Open questions

  • Whether the in vitro selectivity for microbial over mammalian membranes survives at in vivo concentrations and at the doses needed for systemic antimicrobial activity — the central drug-development problem for every cationic amphipathic AMP, not specific to DS5.
  • Whether DS5's anti-Leishmania activity can be translated into a topical or systemic preclinical efficacy signal, given that most in vivo dermaseptin work has focused on dermaseptin S4 analogs rather than DS5 (Krugliak and colleagues, 2000).
  • Whether structural minimization or stabilization (truncation, D-amino acid substitution, lipidation) can preserve the membrane-disruption potency of DS5 while widening its therapeutic window — the medicinal-chemistry program that most of the dermaseptin field has pursued on DS4 has not been extensively replicated on DS5.
  • Whether the synergy reported among native dermaseptins (Mor and colleagues, 1994) reflects a deployable design principle for combination AMP formulations or an artifact of the in vitro assays used to detect it.
Hypotheses5 directions▾ collapse

Research directions for this peptide, selected from the current sources — hypotheses you can explore and model. None of it is proven yet; tap any one to see the full thinking.

openupdated 2026-06-05

Could a single flexible point in the DS5 helix control how deeply the peptide buries itself in bacterial membranes, determining whether it kills Gram-positive or Gram-negative bugs?

If true, chemists could lock or adjust that hinge to make versions of DS5 that specifically target dangerous Gram-negative bacteria like E. coli or Klebsiella, which are increasingly resistant to antibiotics and cause life-threatening hospital infections.

The hypothesis
The glycine at position 17 (G17 in ALWKTLLKKVGTHLAGKAALGAAADKX) acts as a helix-breaking hinge that divides the DS5 amphipathic helix into two independently tiltable segments, and the relative angle between these segments determines the depth of membrane insertion and thus the minimum inhibitory concentration against Gram-negative versus Gram-positive bacteria.
Why it’s plausible
Inspection of the sequence ALWKTLLKKVGTHLAGKAALGAAADKX reveals glycines at positions 11 (G11), 17 (G17), and 21 (G21). In helical peptides, glycine introduces conformational flexibility. The A-L-W-K-T-L-L-K-K-V segment (1-10) is strongly amphipathic (alternating hydrophobic/cationic), while AALGAAADKX (18-27) is alanine/glycine-rich and less amphipathic. A Gly-hinge between these zones could produce a kinked helix whose angle optimises for membrane thickness differences: Gram-negative membranes have a thinner outer leaflet, and a bent helix may better span or destabilise the asymmetric outer membrane. This is supported by the broader dermaseptin literature showing that aggregation and orientation of the helix in membranes is concentration-dependent (doi:10.1074/jbc.272.50.31609).
Why it matters
Identifying G17 as a functional hinge would allow substitution of Gly with alpha-aminoisobutyric acid (Aib) to lock the inter-helix angle and systematically test whether Gram-selectivity is geometrically controlled, providing a rational route to pathogen-selective antimicrobials.
Plausibility.65
Novelty.60
Impact.65
Basis · grounding2 papers · 1 computed/note
[1]
sequenceALWKTLLKKVGTHLAGKAALGAAADKX: G11, G17, G21 are potential helix-breaking residues; the N-terminal amphipathic block and C-terminal alanine/glycine-rich region suggest a two-segment helix
[2]
paper
Energy-transfer and vesicle experiments on dermaseptin family members show concentration-dependent membrane orientation and insertion depth
doi: 10.1074/jbc.272.50.31609
[3]
paper
Dermaseptin S4 analog studies show position-specific substitutions alter aggregation and membrane disruption activity
doi: 10.1074/jbc.275.6.4230
openupdated 2026-06-05

Do parasites like Leishmania attract DS5 to their surface more strongly than human cells do, because the parasites display a specific membrane fat on the outside that human cells keep hidden?

If confirmed, this mechanism would point to a broad strategy for designing selective antiparasitic peptides, potentially improving treatment for neglected tropical diseases that affect tens of millions of people in low-income countries with few affordable options.

The hypothesis
Dermaseptin S5 kills protozoan parasites such as Leishmania at concentrations below its haemolytic threshold because protozoan plasma membranes contain phosphatidylserine exposed on the outer leaflet, which serves as an electrostatic anchor that locally concentrates the cationic DS5 helix to a density sufficient for membrane disruption before equivalent surface density is reached on normal erythrocytes.
Why it’s plausible
Apicomplexan and kinetoplastid protozoa constitutively expose phosphatidylserine on their outer membrane leaflet, unlike healthy mammalian erythrocytes where PS is confined to the inner leaflet. The cationic K-rich DS5 (with five lysines in 27 residues) would be selectively drawn to PS-rich surfaces. This phosphatidylserine-targeting mechanism is documented for other cationic AMPs and would explain the documented anti-protozoan activity of the dermaseptin family without requiring specific receptor interactions.
Why it matters
Leishmaniasis affects 12 million people across 98 countries and treatment remains toxic and limited. If DS5's selective toxicity against protozoa is driven by constitutive PS exposure, this would identify a membrane-lipid target that distinguishes parasite from host and could guide development of next-generation anti-leishmanial peptides.
Plausibility.65
Novelty.50
Impact.70
Basis · grounding2 papers · 1 computed/note
[1]
paper
Dermaseptin family literature covering anti-protozoan activity characterisation, including Leishmania species
doi: 10.1016/j.cbpa.2007.03.016
[2]
sequenceALWKTLLKKVGTHLAGKAALGAAADKX: five lysines create high positive charge density consistent with PS-preferring membrane interaction
[3]
paper
Dermaseptin S4 analog studies demonstrate that charge distribution along the helix determines selectivity windows between cell types
doi: 10.1074/jbc.275.6.4230
openupdated 2026-06-05

If the open end of DS5 were extended with a few extra charged building blocks, could the peptide act as a vehicle to smuggle antifungal drugs into resistant Candida cells that would otherwise pump them out?

Candida auris now causes treatment-resistant hospital outbreaks with mortality rates above 30%. A peptide that smuggles drugs past resistance barriers could restore the effectiveness of existing antifungals, giving clinicians a new tool without requiring entirely new drug development.

The hypothesis
Replacing the C-terminal placeholder residue X in DS5's sequence with a cell-penetrating sequence (e.g., three additional arginines) would convert the membrane-disrupting peptide into a carrier that delivers molecular cargo across fungal cell walls, enabling combination antibiotic-peptide conjugates that bypass the efflux-pump resistance mechanisms that have rendered azole antifungals ineffective in Candida auris.
Why it’s plausible
The C-terminal X in ALWKTLLKKVGTHLAGKAALGAAADKX is a placeholder for the native amide cap. Replacing it with a polyarginine extension would preserve the N-terminal amphipathic helix while adding cell-penetrating capacity at the C-terminus. Candida auris acquires azole resistance primarily through efflux pump overexpression. A DS5-azole conjugate that bypasses the membrane before the drug is pumped out would circumvent this mechanism. The structural basis for this approach is established in the peptide-drug conjugate literature (doi:10.1021/acs.joc.5b01878 shows peptide backbone stability can be engineered for protease resistance).
Why it matters
Candida auris is on the WHO critical priority pathogen list with few treatment options. A membrane-disrupting peptide serving as an intracellular drug delivery vehicle would represent a mechanistic bypass of resistance, directly addressing a major global health threat.
Plausibility.45
Novelty.70
Impact.75
Basis · grounding2 papers · 1 computed/note
[1]
sequenceC-terminal X is a structural placeholder, not a pharmacophore, making it an engineering handle for extension without disrupting the N-terminal active helix
[2]
paper
Demonstrates complete protease resistance is achievable in amphipathic peptide scaffolds through systematic modification
doi: 10.1021/acs.joc.5b01878
[3]
paper
Dermaseptin antifungal activity confirmed against Candida species, providing the relevant activity spectrum
doi: 10.1021/bi00187a034
openupdated 2026-06-05

Could DS5 penetrate and disrupt the sticky biofilm that bacteria form on catheters and implants, where ordinary antibiotics cannot reach them?

Biofilm infections on medical implants kill thousands of patients each year and often require surgically removing the device. A peptide coating that prevented biofilm formation could protect patients from this devastating complication without any need for systemic antibiotic treatment.

The hypothesis
Topical dermaseptin S5 could suppress biofilm-associated Staphylococcus epidermidis infections on implanted medical devices because DS5's amphipathic helix disrupts the lipid-rich extracellular matrix of S. epidermidis biofilms at concentrations below those required to kill planktonic cells, reversing the protective effect of the biofilm without systemic exposure.
Why it’s plausible
S. epidermidis biofilm matrix contains poly-N-acetylglucosamine and extracellular lipids that provide a hydrophobic barrier to conventional antibiotics. DS5's amphipathic character, with the A/G-rich C-terminal segment being less bulky than typical membrane-disrupting helices, suggests it could diffuse through biofilm matrix more readily than larger AMPs. The vaccine literature (doi:10.1016/j.vaccine.2018.01.066) documents anti-viral activity of DS4-type dermaseptins, which involves interaction with lipid envelopes analogous to biofilm lipid components.
Why it matters
Medical device-associated biofilm infections affect millions of patients annually, cause most catheter-related bloodstream infections, and are largely untreatable without device removal. A topically applied peptide coating that disrupts biofilm matrix without systemic toxicity could dramatically reduce implant infection rates.
Plausibility.55
Novelty.50
Impact.70
Basis · grounding2 papers · 1 computed/note
[1]
paper
Dermaseptins S3, S4, S4M4K show antiviral activity involving lipid envelope disruption, demonstrating that the helix acts on complex lipid structures beyond simple bilayers
doi: 10.1016/j.vaccine.2018.01.066
[2]
paper
Demonstrates that dermaseptin-class peptide scaffolds can be made fully protease-resistant, a prerequisite for stability in a device coating environment
doi: 10.1021/acs.joc.5b01878
[3]
sequenceALWKTLLKKVGTHLAGKAALGAAADKX: alanine/glycine-rich C-terminus is less bulky than typical beta-hairpin AMPs, suggesting better biofilm penetration potential
openupdated 2026-06-05

Does the unusual number and spacing of positively charged amino acids in DS5 help the peptide target the type of membrane fat found in fungi but not in human cells?

Fungal infections kill over 1.5 million people per year, and options are limited. If DS5 can selectively target fungal membranes, derivatives could provide a new class of topical or wound antifungals with less risk of harming human tissue.

The hypothesis
Dermaseptin S5's high lysine content (K4, K8, K9, K17, K26 in the sequence ALWKTLLKKVGTHLAGKAALGAAADKX) confers selective toxicity toward fungal over mammalian membranes because fungal membranes are enriched in ergosterol, which packs less tightly than cholesterol, and the resulting reduced membrane order allows DS5 to insert at lower peptide concentrations than required for cholesterol-containing mammalian membranes.
Why it’s plausible
Five lysines distributed along the 27-residue sequence create a high local positive charge density. Ergosterol-rich fungal membranes have a lower dipole potential and looser lipid packing than cholesterol-rich mammalian membranes. Cationic peptides with distributed charge can exploit this differential by inserting preferentially into less ordered bilayers at sub-haemolytic concentrations. This selectivity window explains the anti-fungal activity described in Mor et al. 1994 (doi:10.1021/bi00187a034) while the peptide retains cytotoxicity at higher concentrations.
Why it matters
Fungal infections are the fourth leading cause of hospital-acquired infection and are increasingly drug-resistant. If DS5's selectivity for ergosterol-membrane geometry can be quantified and enhanced through conservative lysine-spacing modifications, it could become a lead for topical or wound-care antifungals.
Plausibility.60
Novelty.40
Impact.65
Basis · grounding2 papers · 1 computed/note
[1]
sequenceALWKTLLKKVGTHLAGKAALGAAADKX: K at positions 4, 8, 9, 17, 26 gives approximately one lysine per 5 residues, high positive charge density
[2]
paper
Mor et al. original dermaseptin isolation paper characterises antifungal activity using antifungal assays as a purification guide, confirming anti-Candida activity
doi: 10.1021/bi00187a034
[3]
paper
Fluorescence-based membrane binding studies of dermaseptin family show lipid-composition-dependent insertion kinetics
doi: 10.1074/jbc.272.50.31609
details expand to inspect
3-letter notation
Ala-Leu-Trp-Lys-Thr-Leu-Leu-Lys-Lys-Val-Gly-Thr-His-Leu-Ala-Gly-Lys-Ala-Ala-Leu-Gly-Ala-Ala-Ala-Asp-Lys-X
citationbibtex
peptidemodel (2026). Dermaseptin S5: frog-skin natural germ-killer (DS5) (pep-10966, v1). PeptideModel. https://peptidemodel.com/card/pep-10966
@peptide{pep10966,
  sequence = {ALWKTLLKKVGTHLAGKAALGAAADKX},
  target   = {},
  author   = {peptidemodel},
  year     = {2026},
  status   = {designed}
}
clinical trials 0 trials · checked 2026-05-22
0
no registered clinical trials as of 2026-05-22; we'll re-check periodically
references 12 papers
[1] supporting
[2]
Antimalarial Activities of Dermaseptin S4 Derivatives
Krugliak, M. et al. Antimicrobial Agents and Chemotherapy 2000
supporting
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